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Acknowledgements We thank Anna Altin, Tamara Grossrieder, Arne Hansen, Samanda Krasniqi, Marina Reist, Melanie Schulz and Jelena Velimirovic, for their help in data acquisition, and Samuel Gerster for technical support

The consequent sequestration of redox-active Cu in the extracellular compartment and certain intracellular pools is a key mediator of cytotoxicity and cardiotoxicity in diabetes [51, 170]